Titus was born on 24 August 1974. He was named by gorilla researcher Dian Fossey, who was observing his group at the time, after the eponymous protagonist of the Mervyn Peake novel Titus Groan. When Titus was four years old a blackback male joined his group and was named 'Beetsme' by Fossey
Titus appears to have formed a bond with Beetsme and the pair became the nucleus of an unstable bachelor group. The group was without females for eight years, during which time they were observed engaging in homosexual behaviour.
What the what! However, it should be noted that Titus eventually overcame his gayness to sire "more children than any other gorilla on record". Although he may have had to fantasize about male gorillas to perform the actual penetrations - science isn't sure. But it's still an inspirational story! And now Titus is dead!